Set in 1993, the pair are both together with their owners as they are about to move to a new home. The moving van is at their old house waiting, and Tom doses in the back of the car. However when he notices Jerry, he puts him on a stick, and Jerry, noticing no escape and knowing that he will fly, he grabs hold of Tom's whiskers so they fly together into the garden. Jerry quickly dashes into his mousehole and locks the door, Tom nailing wooden planks on the door. Tom leaves the house, but it's too late: The owners have left. When Tom tries to get in the moving car, he ends up with a bulldog and ties up his ears so he cannot see. Tom runs into the house for safety and stays there for the night. The next day, Tom notices that the house is being destroyed by a demolition crew. He manages to escape but realizes he couldn't just leave Jerry goes back and saves him, the two manage to survive. But now homeless, they meet a dog named Puggsy and his friend Frankie Da Flea that tries to teach Tom and Jerry to be friends. Then they all agree to have a 'feast' at their place and Puggsy makes a 'buffet' by collecting leftovers in the bin. When Puggsy's tray is crammed, a dogcatcher captures Puggsy and Frankie and lock them in their car.
Tom and Jerry then meet an eight-year old girl named Robyn Starling, daughter of Daddy Starling, whose mother died of pneumonia when she was a baby and is left behind with her horrid and evil guardian Aunt Figg when her dad goes away to Tibet. Robyn runs away after her locket is thrown out the window and that's how she began to run. Jerry said to her that if she runs, her things won't be with her but Robyn said to them that Aunt Figg may seem sweet but she's real mean.
Cut to Aunt Figg crying in the house, scared of losing Robyn. With the help of her evil lawyer, Lickboot and her overweight dachshund, Ferdinand, make a reward of a million dollars for the return of Robyn, who they wish to sell for a ransom, engulfed by love of money. Robyn is recaptured, but manages to escape yet again, after Tom and Jerry are kidnapped by Dr. Applecheek and are sent to a city pound where animals are abused. At this point, everybody is looking for the million dollar girl, and Aunt Figg and Lickboot manage to get to Robyn's escape destination first. What was planned as another capture and it only worsens the situation when an oil lamp is knocked on the floor. As the house goes up in flames, Robin's father finally arrives and rescues them. Tom and Jerry are taken to a new home where they both promise not to trick each other ever again. But the pair have soon reverted back to their old ways, and the movie finishes with a fun, loving classic scene of Tom chasing Jerry into the distance...or just in their new home.
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